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Mitochondrial dysfunction from any cause (biotoxin illness, heavy metals, oxidative stress) directly impairs amino acid conjugation because the activation step requires mitochondrial ATP and CoA

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(PubMed) Diez-Ricote L, San-Cristobal R, Concejo MJ, et al

In response to agonist stimulation, GPCRs are phosphorylated by GRKs, after which GPCRs promote arrestin recruitment to the activated receptors

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Glutathione shifts the process of melanogenesis from eumelanin to pheomelanin through the reaction between thiol and dopaquinone groups causing formation of sulfhydryl-dopa conjugates[] [Figure 2]