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Data Availability RNA sequencing data are accessible online at Mendeley Data ( Acknowledgements We thank Tim Stearns for help with confocal imaging, Edo Biluar for reading the manuscript and assisting with experiments, and Julien Sage and Jan Carette for providing reagents

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Alterations in calcium homeostasis can lead to mitochondrial swelling (Mller et al., 2011), resulting in the leakage of Ca 2+ ions from mitochondria