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Upon reaching 90% confluency, cells were rinsed with Dulbeccos phosphate-buffered saline (DPBS) without calcium and magnesium then detached using 1.5 mL of 0.25% trypsin EDTA/DPBS with polyvinylpyrrolidone for 1.5 min at 37 C

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As described in greater detail elsewhere [30] the coverage and completeness of these datasets varied, therefore a novel approach to filling these gaps and ensuring the temporal validity of data was developed using Google Street View images